Funny movie...as usual, Betty White and Kristian Chenoweth are the best!
The film opens in 2002 with Marni (Kristen Bell), an acne-riddled high school senior at Ridgefield High School, with glasses and braces, making a video about how much she hates high school, and reveals how she is tormented and bullied by other girls, specifically Joanna (Odette Yustman), who made Marni's high school life miserable. She adds that her protective older brother, Will, (James Wolk), was very popular as a handsome basketball player. However, at a very important basketball game, Joanna pushes the mascot (Marni), who runs into Will, resulting in a loss of the game.
Years later, in 2010, however, she is a successful public relations executive, recently promoted to a job in New York. When she flies home to her older brother's wedding, she discovers her brother Will is about to marry Joanna, who bullied and tormented her throughout high school. When Marni meets Joanna for the first time in eight years, Joanna seems to not recognize her. Marni is also upset to see that Joanna fits in very well with the family. The plot thickens when her mother Gail (Jamie Lee Curtis) meets up with Joanna's aunt Ramona (Sigourney Weaver), Gail's former best friend who pushed her into a pool at their senior prom. Ramona is now a successful, wealthy woman who owns several hotels and a private plane.
Although Gail seems willing to put the past behind her, she still feels the need to "outdo" Ramona during their interactions together. On the other hand, Marni is unwilling to forget the things Joanna did to her in high school, unless she apologizes, and decides to try to let her brother know of Joanna's bullying past. Her attempts to get Will alone for a conversation fail. When Marni tells Joanna that she knows who she really is, it is obvious that Joanna remembers Marni. She refuses to give Marni a meaningful apology, and treats her disrespectfully, leaving Marni convinced that Joanna has not changed. Meanwhile, Gail comes to Ramona's hotel room for "closure" about anything bad between them that happened in the past. They appear to make up, but Ramona still seems not to be too fond of Gail.
One day on the street, Will and Marni's grandmother Bunny (Betty White) meets a man named Tim (Kyle Bornheimer). Joanna reveals that they have dated, and Tim appears devastated when he hears that she's marrying Will. Marni decides to bring Tim to the rehearsal dinner as part of her plan to stop the wedding. When it is time for guests to make a toast to the bride and groom, Tim unexpectedly jumps up to give his toast to Joanna. He reveals to a stunned wedding party that Joanna left him at the altar. Later, a video is presented (recovered by Marni) from their old high school time capsule. The video reveals Joanna as she was in high school: an alpha-female bully, with the footage showing proof of her tormenting Marni and pushing other students. Will unplugs the video projector before the video is complete and walks out, furious.
However, Marni is in trouble when everyone discovers that she was responsible for the video at the rehearsal dinner. Marni decides to talk with Joanna, hoping to reconcile, but Joanna is too furious and tells Marni "I thought we could start over." Marni is now convinced that Joanna hasn't changed, and walks away. Joanna then starts a fight with Marni, who fights back. Will walks in and witnesses the fight. He tells Joanna that he doesn't care about her past, but is angry that she was not being honest with him. He then tells Marni that he is furious at her for going behind his back.
Meanwhile, Ramona and Gail argue after the rehearsal dinner, and Ramona accuses Gail of trying to ruin her life throughout high school. A catfight ensues, with both of them falling into the pool. Ramona reveals that she had a grudge with Gail, even when they were best friends, because she always competed with and outdid her, culminating with Gail taking the boy that Ramona wanted to the prom. Ramona stated that Gail was already a legendary head cheerleader and prom queen, and drama was supposed to be hers, yet Gail also defeated her with the auditions. She sarcastically thanks Gail for what she did, because it motivated her to become successful in life. Gail apologizes for being insensitive, but reveals that she is proud of Ramona, and that her loving family is her accomplishment. Ramona, remorseful of her actions, tells Gail that she was jealous of Gail's happy family especially that her marriages didn't work out, and feared that she was trying to take Joanna away from her. The two reconcile their friendship.
Later that night, Marni finds Joanna in the kitchen binging on junk food. She finally admits to Marni that she feels truly awful for bullying and tormenting her and feels like an awful person, and that she loves Will. Marni forgives her and promises to get them back together. The next day,during a jog, Marni apologizes to Will for her actions, saying she was only trying to protect him.
Joanna and Will reconcile in the family's old tree house, but it collapses and injures both of them when Ben Olsen (Billy Unger) loosened the screws as a part of his plan in hiding the tree house. They are both forced to stay at a hospital, which delays the wedding. However, Marni puts together a makeshift wedding at the hospital, with the bride and groom bandaged, but properly dressed and able to walk down the aisle. Marni appears to start a relationship with Charlie (Sean Wing), her brother's best friend who was always kind to her. Joanna introduces Marni's grandmother Bunny to Helen Sullivan (Cloris Leachman). It is revealed that Helen and Bunny were enemies in high school when Helen stole a boy from her. However, Bunny gets her revenge when she cuts in on Helen's dance and takes her partner.
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